Asexual reproduction is the production of progeny without the union of cells or nuclear material. Many small algae reproduce asexually by ordinary or by fragmentation, whereas larger algae reproduce. Some red algae produce monospores (walled, nonflagellate, spherical cells) that are carried by water currents and upon produce a new organism. Some produce nonmotile spores called, while others produce, which lack true cell walls and bear one or more. These flagella allow zoospores to swim to a favourable, whereas monospores and aplanospores have to rely on passive transport by water currents.is characterized by the process of, in which progeny cells receive half of their genetic information from each parent cell.
Sexual reproduction is usually regulated by environmental events. In many, when temperature, salinity, inorganic nutrients (e.g., and ), or day length become unfavourable, sexual reproduction is induced. A sexually reproducing organism typically has two phases in its life cycle. In the first stage, each cell has a single set of and is called, whereas in the second stage each cell has two sets of chromosomes and is called. When one haploid fuses with another haploid gamete during fertilization, the resulting combination, with two sets of chromosomes, is called a. Either immediately or at some later time, a diploid cell directly or indirectly undergoes a special reductive cell-division process (meiosis).
The red algal life history is typically an alternation of generations that may have three generations rather than two. Chloroplasts evolved following an endosymbiotic event between an ancestral, photosynthetic cyanobacterium and an early eukarytoic phagotroph.
Diploid cells in this stage are called because they produce spores. During meiosis the of a diploid is halved, and the resulting daughter cells are haploid.
At some time, immediately or later, haploid cells act directly as gametes. In algae, as in plants, haploid cells in this stage are called because they produce gametes. Most algae can live for days, weeks, or months. Small algae are sometimes found in abundance during a short period of the year and remain dormant during the rest of the year.
In some species, the dormant form is a resistant cyst, whereas other species remain in the vegetative state but at very low population numbers. Some large, attached species are true. They may lose the main body at the end of the, but the attachment part, the holdfast, produces new growth only at the beginning of the next growing season.The, as exemplified by, have some of the most complex life cycles known for living organisms. Following meiosis, four haploid are produced, which germinate to produce either a male or a female gametophyte.
When mature, the male gametophyte produces special branches that bear structures, called spermatangia, which contain spermatia, the male gametes. The female gametophyte produces special branches that bear carpogonia, the female gametes. Fertilization occurs when a male spermatium, carried by water currents, collides with the extended portion of a female carpogonium and the two gametes fuse. The fertilized carpogonium (the zygote) and the female gametophyte around it develop into a basketlike or pustulelike structure called a carposporophyte. The carposporophyte eventually produces and releases diploid carpospores that develop into tetrasporophytes. Certain cells of the tetrasporophyte undergo meiosis to produce tetraspores, and the cycle is repeated.
In the life cycle of Polysiphonia, and many other red algae, there are separate male and female gametophytes, carposporophytes that develop on the female gametophytes, and separate tetrasporophytes.The life cycles of, which are diploid, are also unique. Diatom walls, or frustules, are composed of two overlapping parts (the valves). During cell division, two new valves form in the middle of the cell and partition the protoplasm into two parts. Consequently, the new valves are generally somewhat smaller than the originals, so after many successive generations, most of the cells in the growing population are smaller than their parents. When such diatoms reach a critically small size, sexual reproduction may be stimulated. The small diploid cells undergo meiosis, and among pennate (thin, elliptical) diatoms the resulting haploid gametes fuse into a zygote, which grows quite large and forms a special kind of cell called an auxospore.
The auxospore divides, forming two large, vegetative cells, and in this manner the larger size is renewed. In centric diatoms there is marked differentiation between nonmotile female gametes, which act as egg cells, and motile (typically uniflagellate) male gametes.